Spogostylum niphasoides sp. nov.
Description
Male (holotype) (Fig. 2). Relatively medium to large-sized robust species, body length is about 13 mm and wing length is about 13.2 mm. Head. Black in ground color; frons white dusted, with upper frons narrowed, about two times as wide as ocellar tubercle, all frons covered with pure white hairs, becoming slightly yellowish at upper frons; face densely covered with shaggy long pure white hairs; antennae 1 mm in length, yellowish-brown to brown at upper half of scape, lower half of pedicel and narrow apical part of flagellum, dark brown at broad lower base of flagellum, upper half of pedicel and lower half of scape; lower base of flagellum dusted white; hairs of antennae long white on scape, relatively short white on pedicel mixed with few black ones at outer side; scape about two times as long as and slightly wider than pedicel. Thorax. Glossy black in ground color, covered with shaggy long white hairs and fine short hairs, becoming slightly yellowish at central parts of scutum and scutellum; bristles of scutum and scutellum white, except yellowish brown supra-alar bristles; wings hyaline with very faint yellowish basicostal infuscation and whitish costal cell (c); veins dark brown except yellowish subcostal vein (Sc) and humeral cross-vein (h); base of costal vein white dusted, with short black hairs; costal vein (C) with shorter black hairs along margin; stem vein white dusted; anal cell (cup) narrowly open at wing margin, with opening about half length of posterior cross-vein (bm-cu) [in S. niphas, opening of cup relatively narrower, about third length of bm-cu or less]; halters yellowish, dusted white especially at tip of knob; legs yellowish-brown except the last tarsomeres of fore and middle legs, and all tarsomeres of hind legs which are brown; all legs covered with white scales and fine hairs, with short black bristles, bristles on tibia about half tibial width in length. Abdomen. Tergites glossy black in ground color with reddish-brown posterior borders, covered with white scaly hairs and long silky white hairs, becoming more dense and longer at sides and posterior margins, forming transverse bands almost on all tergites; some yellowish to brown fine hairs, and scaly hairs may also present mixed with white scaly hairs of white bands at margins of tergites and on center of second tergite; posterior tergite densely covered with broader scales and longer silky white hairs; gonocoxites truncate, triangular; gonostyli broad at base, with long curved apical part, constricted between broad base and long apical part, with small outer swelling covered with long hairs; aedeagal sheath is slightly longer than and enclosing aedeagus, with a broad complicated tip having two thorn-like dark spines directed antero-dorsally and located at proximal border of tip complex, and two smaller spines dorsally approximately at middle of tip complex; from lateral view, tip complex of aedeagal sheath seems semi-quadrangular, more sclerotized and darker in color dorsally, less sclerotized and lighter in color ventrally.
Female unknown.
Material examined. Holotype ♂, Wadi Garagneyia (South Sinai), VI–VIII.42 (Efflatoun); paratype ♂ (in a poor condition), same data as holotype; all preserved in EFC.
An updated checklist of Spogostylum spp. in Egypt
Spogostylum bisniphas (Bezzi, 1925)
Distribution: PA: Egypt.
Egyptian localities: Eastern Desert: Wadi Dar El-Maskhara, Wadi Garawi, Wadi Hoff, Wadi Rishrash, Wadi Um-Girfan.
Dates of collection: April to June.
Spogostylum candidum (Sack, 1909)
Distribution: AF: Egypt [as “Gebel Elba”], United Arab Emirates. OR: Pakistan. PA: Egypt, Iran, Saudi Arabia, Turkey.
Egyptian localities: Eastern Desert: Abu-Sueir, Geneifa, Wadi Garawi, Wadi Murrah; Sinai: El-Arish, Bir Tarfah, Gebel Moussa, Hagar Bardeyia, Rafah, Wadi El-Arbaein, Wadi El-Rabba, Wadi Firan, Wadi Garagneyia, Wadi Isla, El-Zawatein; Fayoum; Gebel Elba.
Dates of collection: March to August.
Spogostylum efflatouni Paramonov, 1957
Spongostylum sergeiae Hull, 1973.
Distribution: PA: Egypt.
Egyptian localities: Eastern Desert: Wadi Digla.
Dates of collection: April to June.
Spogostylum griseipenne (Macquart, 1850)
Distribution: AF: Mauritania, Niger, United Arab Emirates. PA: Egypt, Iraq, Israel, Libya, Palestine (West Bank), Saudi Arabia, Tunisia.
Egyptian localities: Lower Nile Valley: Ebn Salam (Mansourah);Western Desert: Kharga; Fayoum: Kom Osheem.
Dates of collection: April to November.
Spogostylum hamadnallahi El-Hawagry, 2002
Distribution: PA: Egypt.
Egyptian localities: Sinai: Wadi Garagneyia.
Dates of collection: August.
Spogostylum hippolyta (Wiedemann, 1828)
Argyromoeba dedecor Hermann, 1907.
Anthrax villosa Klug in Hull, 1973. Nomen nudum.
Distribution: AF: Eritrea, Gambia, Guinea, Mali, Mauritania, Senegal, Sudan. PA: Egypt, Iran, Israel, Libya, Tajikistan, Turkey, Turkmenistan.
Egyptian localities: Coastal Strip: Hammam; Western Desert: Kharga; Sinai: Moweilah.
Dates of collection: September.
Spogostylum incisurale (Macquart, 1840)
Anthrax biappendiculata Macquart, 1855.
Anthrax mixta Loew, 1860.
Molybdamoeba incisuralis var. inquirenda Bezzi, 1912.
Argyramoeba incisuralis f. aridicola Hesse, 1956.
Argyramoeba incisuralis F. glaucescens Hesse, 1956.
Argyramoeba incisuralis f. fumosa Hesse, 1956.
Distribution: AF: Botswana, Egypt [as “Gebel Elba”], Lesotho, Malawi, Mozambique, Namibia, South Africa, Tanzania, Uganda, Zambia, Zimbabwe. PA: Egypt.
Egyptian localities: Coastal Strip: Alexandria, Nuzha, Ramleh; Lower Nile Valley: Abu-Rawash, Barrage, Borgash, Faraskour, KafrHakim, Katta, Kirdassa, Mansouriah, Ezbet Nakhl; Eastern Desert: Gebel Asfar, Ismailia, Wadi Garawi, Wadi Um-Elik; Fayoum: Kom Osheem; Sinai: Wadi El-Lega, Wadi Firan; Gebel Elba.
Dates of collection: April to September.
Spogostylum isis (Meigen, 1820)
Anthrax subnotata Walker, 1871.
Distribution: PA: Algeria, Armenia, Austria, Azerbaijan, Belgium, Croatia, Cyprus, Egypt, France, Greece, Gruzia, Israel, Italy, Kazakhstan, Kyrgyz Republic, Libya, Macedonia, Malta, Morocco, Portugal, Saudi Arabia, Spain, Syria, Tajikistan, Tunisia, Turkey, Turkmenistan, Uzbekistan, Yugoslavia.
Egyptian localities: Coastal Strip: Mariout; Lower Nile Valley: Kafr Hakim, Kirdassa; Eastern Desert: Abu-Sueir, Fayed, Geneifa, Wadi Gharagid, Wadi Ibtadi, Wadi Um-Girfan, Wadi Zohleiga; Sinai: Wadi El-Arbaein, Wadi Isla, Wadi Tlah.
Dates of collection: April to August.
Spogostylum niphas (Hermann, 1907)
Spongostylum niphas var. bilineata Engel, 1936.
Distribution: AF: Eritrea, Mauritania, Oman, Somalia, Yemen. OR: Pakistan.PA: Egypt, Israel, Saudi Arabia, Tajikistan, Turkey, Turkmenistan.
Egyptian localities: Coastal Strip: Mariout; Lower Nile Valley: Barrage, Helwan, Mansouriah, Tisfa; Eastern Desert: Wadi Garawi, Wadi Girfan, Wadi Ibtadi.
Dates of collection: April to August.
Spogostylum niphasoides
sp. nov.
Distribution: PA: Egypt.
Egyptian localities: Sinai: Wadi Garagneyia (South Sinai).
Dates of collection: June to August.
Spogostylum ocyale (Wiedemann, 1828)
Anthrax umbra Walker, 1849.
Spogostylum arugotense Zaitzev, 1997.
Distribution: AF: Oman, Somalia, Sudan, United Arab Emirates, Yemen. PA: Egypt, Iran, Israel, Jordan, Libya, Palestine (West Bank), Saudi Arabia.
Egyptian localities: Coastal Strip: Amriah, Matrouh, Mex, Nuzha, Salloum; Lower Nile Valley: Barrage, Borgash, Helwan, Kirdassa; Fayoum; Eastern Desert: Gubbet El-Bous, Wadi Digla, Wadi Hoff; Western Desert: Dakhla; Upper Nile Valley: Abu-Simbel, El-Minya; Sinai: El-Arish, Wadi Firan, Wadi Isla, Wadi Moueilah.
Dates of collection: January to October.
Spogostylum sordidum Sack, 1909.
Spongostylum antiopa Bezzi, 1925.
Distribution: AF: Oman. PA: Afghanistan, Egypt, Iran, Saudi Arabia, Tajikistan, Turkmenistan, Uzbekistan.
Egyptian localities: Coastal Strip: Mariout, Matrouh, Nuzha, Salloum; Eastern Desert: Wadi Digla, Wadi Hoff; Sinai: El-Arish.
Dates of collection: January to September.
Spogostylum tripunctatum (Pallas in Wiedemann 1818)
Anthrax difficilis Wiedemann in Meigen, 1820.
Spogostylum mystaceum Macquart, 1840.
Distribution: PA: Afghanistan, Albania, Algeria, Armenia, Austria, Azerbaijan, Bosnia-Herzegovina, Bulgaria, Croatia, Egypt, France, Greece (incl. Ithaca), Gruzia, Israel, Italy, Kyrgyz Republic, Libya, Macedonia, Malta, Moldova, Morocco, Russia, Spain, Switzerland, Syria, Tajikistan, Turkey, Turkmenistan, Ukraine, Uzbekistan.
Egyptian localities: Lower Nile Valley: Ezbet El-Nakhl, Mansouriah; Fayoum.
Dates of collection: March to June.
Spogostylum ventrale Bezzi, 1924
Distribution: AF: Eritrea, Ethiopia, Kenya, Oman, Yemen. PA: Egypt.
Egyptian localities: Sinai: Wadi El-Arbaein.
Dates of collection: May to July.
Spogostylum volitans Wiedemann, 1828
Distribution: AF: Sudan. PA: Egypt.
Egyptian localities: unknown.
Dates of collection: unknown.