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Table 1 Functional response of H. axyridis on C. juglandicola and P. juglandis under four different temperature regimes

From: Effect of temperature on predation by Harmonia axyridis (Pall.) (Coleoptera: Coccinellidae) on the walnut aphids Chromaphis juglandicola Kalt. and Panaphis juglandis (Goeze)

Aphids

Temp. (°C)

Attacking efficiency (a)

Handling time Th (d)

Predation capacity a/Th

Maximum predation number 1/Th

Holling’s type II equation

R2

C. juglandicola

15

0.7944

0.0031

256.2604

322.5806

Na = 0.7944N/(1 + 0.0025N)

0.8756

20

0.9169

0.0021

434.5543

473.9336

Na = 0.9169N/(1 + 0.0019N)

0.9821

25

1.0158

0.0020

495.5050

487.8049

Na = 1.0158N/(1 + 0.0021N)

0.9964

30

1.0862

0.0019

565.7481

520.8333

Na = 1.0862N/(1 + 0.0021N)

0.9923

P. juglandis

15

0.5798

0.0075

77.2065

133.1558

Na = 0.5798N/(1 + 0.0044N)

0.9641

20

0.8034

0.0042

189.4811

235.8491

Na = 0.8034N/(1 + 0.0034N)

0.9969

25

0.8386

0.0035

236.8973

282.4859

Na = 0.8386N/(1 + 0.0030N)

0.9978

30

0.8468

0.0029

293.0194

346.0208

Na = 0.8468N/(1 + 0.0024N)

0.9923